Pretty Little Liars: Before They Were Stars

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Ashley Benson: Then

Signed by the Ford Modeling Agency at age eight, Benson appeared in commercials and small TV roles before taking on the role of Abigail Deveraux on Days of Our Lives in 2004. She had her first kiss -- on TV -- with DOOL costar Brock Kelly at age 15.

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Ashley Benson: Now

Benson appears as Hanna Marin on Pretty Little Liars.

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Lucy Hale: Then

Born in 1989 in Memphis, Hale got her big break in 2003 as a contestant on FOX's singing competition, American Juniors, a precursor to American Idol also hosted by Ryan Seacrest.

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Lucy Hale: Now

Hale appears as Aria Montgomery on Pretty Little Liars.

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Troian Belliario: Then

Assuming small roles in her TV producer dad Donald P. Bellisario's shows Quantum Leap and NCIS, Bellisario appeared alongside Mary-Kate and Ashley Olsen in the film Billboard Dad. "When I was 5 years old, my best friends were Mary-Kate and Ashley," she tells Us. "I just knew them as the girls across the street!"

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Troian Bellisario: Now

Bellisario appears as Spencer Hastings on Pretty Little Liars.

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Shay Mitchell: Then

Before PLL, the Canadian native earned cash as a model. "I did it for six years, and lived in Thailand and Hong Kong. It was fun!" she tells Us of her former gig.

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Shay Mitchell: Now

Mitchell appears as Emily Fields on Pretty Little Liars.

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Ian Harding: Then

Growing up as an army brat in both Heidelberg, Germany and Virginia, Harding joined the drama club at his Maryland high school and scored his first film role, as Ryan in Deadtime Stories 2, in 2009. He also appeared in Adventureland that same year.

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Ian Harding: Now

Harding appears as Ezra Fitz on Pretty Little Liars.

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Laura Leighton: Then

Leighton's first major acting role was as Sydney Andrews in TV's Melrose Place, on which she appeared from 1993-1997. Leighton (photographed in 1998) is married to her Melrose Place costar Doug Savant.

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Laura Leighton: Now

Leighton appears as Ashley Marin on Pretty Little Liars.

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Keegan Allen: Then

"I took Shakespeare classes and carried my camera around in high school," the lifelong Los Angeleno tells Us of growing up pre-PLL.

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Keegan Allen: Now

Allen appears as Toby Cavanaugh on Pretty Little Liars.

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Janel Parrish: Then

Hawaiian-born Parrish began her career as a member of the National Touring Company and Broadway Company of Les Miserables as Little Cosette. Later performing in the all-girl group Impulse, Parrish took on a role in the 2007 Bratz movie.

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Janel Parrish: Now

Parrish appears as Mona Vanderwaal on Pretty Little Liars.

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Tyler Blackburn: Then

Also a musician, Blackburn starred as Nathan on Nickelodeon's Unfabulous in 2005. Bit roles in the films The Doers of Coming Deeds and Next of Kin followed.

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Tyler Blackburn: Now

Blackburn appears as Caleb Rivers on Pretty Little Liars. He'll also star in the PLL spinoff Ravenswood.

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Sasha Pieterse: Then

Pieterse made her acting debut at age 6 in 2002 as Buffy in TV's Family Affair. Three years later, the actress made her film debut in The Adventures of Shark Boy and Lava Girl and found her guest appearance on House M.D. nominated for a daytime Emmy.

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Sasha Pieterse: Now

Pieterse appears as Alison DiLaurentis on Pretty Little Liars.