FIRST PIC: Jennifer Aniston Kisses Justin Theroux!

 London Entertainment/Splash

Wednesday was a day of milestones for Jennifer Aniston.

In an emotional ceremony, the actress, 43, received her star on Hollywood's Walk of Fame as friends, family and fans looked on. To celebrate the moment, she gave beau Justin Theroux a kiss on the lips.

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Although she and her sexy Wanderlust costar, 40, went public with their romance almost a year ago, it's the first time they've been photographed in a lip lock.  During the ceremony — with speeches by Adam Sandler and her Wanderlust costars Malin Ackerman and Kathryn Hahn — Theroux proudly looked on, sitting next to his would-be father-in-law, Days of Our Lives star John Aniston. See more pics from Jennifer's big day.

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"I cannot explain to you how surreal this moment is!" Aniston gushed at the podium. "This is crazy! I mean, literally. I've walked the streets of Hollywood Boulevard," she said before catching the double entendre.  "Well, no no no, oh please. I didn't mean like that. You know. I think we're about to get on all fours too, so this could get really interesting!"

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Tearfully, she added, "I am a California girl through and through, and this has always been in the back of my mind. It was one of my wildest dreams . . .I am utterly humbled and grateful. . . and to all of the fans who have come out today and supported me throughout my career, my family, my papa, and my dear friends. . . I LOVE YOU!"

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