See how Katie Holmes, James Van Der Beek, Michelle Williams and Joshua Jackson have grown since being cast on The WB series in 1998
Joshua Jackson - 2012
Since 2008, Jackson has played jack-of-all-trades Peter Bishop on the FOX hit Fringe. His character later became romantically involved with Olivia Dunham (Anna Torv). He'll next appear on the big screen in Lay the Favorite alongside Bruce Willis, Catherine Zeta-Jones, Vince Vaughn and Laura Prepon.
Credit: Liane Hentscher/FOX