The O.C. Cast: Then and Now

Rachel Bilson: Then

Bilson's Summer Roberts was originally intended to appear in just a few episodes, but the actress charmed audiences and was soon upgraded to series regular. Throughout the show's four seasons, her character was involved in an on-and-off relationship with the adorkable Seth Cohen (Adam Brody, whom she also dated in real life). In the series finale, Summer and Seth tied the knot.

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